Key to the genera of Tachinidae

in the eastern US

Fourth page of the Tachinid key. See page one for information and initial key points. Eastern species list.

Morphology site of O'Hara

197.   Eye with short sparse hairs; each hair no longer than combined diameter of two or three facets


Goniocera B. & B. (1/1)

io Aldrich



197.   Eye densely covered with conspicuous long hairs







197A.   Pedicel with dense patch of stout erect bristles, each about as long as width of pedicel at point of attachment. Longest parafacial hairs more than half as long as arista


M. tibialis


Metopomuscopteryx Townsend (1/3)

tibialis Coquillett



197A.   Pedicel with few, sparse hairs and one or tow bristles, most shorter than width of pedicel. Parafacial hairs at most less than one-third as long as arista


M. obscura


Macquartia R-D. (2/7)




198.   Scutum without bristles except for one pair of each of the following: presutural supra-alar, posteriormost presutural, postsutural supra-alar, postsutural intra-alar, postsutural dorsocentral, and postsutural acrostichal; usually only one katepisternal bristle present, in posterodorsal corner (below). Abdominal tergites without dorsal bristles






 198.   Scutum with two or more pair of postsutural supra-alar, dorsocentral, and acrostichal bristles; katepisternum with two or more bristles (below). Abdominal tergites usually with at least one pair of marginal bristles








199.   Hind tibia with dorsal longitudinal row of long flattened blade-like setae, each as long as width of tibia at its point of attachment, flanked by a second row of similarly shaped but much shorter setae



Trichopoda Berthold (5/6 in 2 subgenera)



199.   Hind tibia lacking rows of long flattened setae







200.   Palpus minute, much shorter than pedicel. Abdomen orange-red without dark markings, convex dorsally, twice as long as wide, and covered with prominent recumbent black hairs, none of which is enlarged or bristle-like; apex of abdomen of female folded ventrally; ventral edges of tergite 2 studded with spine-like setae



                                                                        B. brevipennis


Besseria R-D., in part (2/3)



200.   Palpus well-developed, longer than pedicel and usually longer than antenna. Abdomen, if orange-red, with dark markings; apex of abdomen of female not folded ventrally and not opposed by spine-like setae on ventral side of tergite 2







201.   Abdomen flattened dorsally, distinctly longer than wide, usually dark colored, but if pale in ground color, then densely pruinose


Phasia Latreille (10/16)



201.   Abdomen hemispherical dorsally, scarcely longer than wide, orange red in ground color, usually with dorsal dark markings, sometimes almost black in female







202.   Apex of scutellum without bristles; apical pair so widely separated and short that they appear as lateral bristles and cannot cross at their apices. Abdomen shiny, orange red with black markings, or sometimes entirely black, with short hairs each arising from a raised base, giving abdomen a somewhat rugose appearance. Pedicel and first flagellomere in most species elongate, extending to level of vibrissa


G. canadensis


Gymnosoma Meigen (5/6)



 202.   Apex of scutellum with the usual crossed pair of apical bristles. Abdomen usually golden tomentose (in male) or gray tomentose (in female); intertergal sutures distinct. Pedicel and first flagellomere short, not extending to level of vibrissa



                                                                    G. immaculata


Gymnoclytia B. & B. (5/6)




203.   Postmetacoxal area sclerotized, i.e. posterior edges of metepimera extending posteromedially to meet each other at midline between hind coxae and base of first abdominal sternite, forming a postmetacoxal bridge






 203.   Postmetacoxal area membranous, distinctly paler in color than adjacent metepimera







204.   Hind margin of eye indented. Ocellar triangle extending forward to lunule as a polished brown strap-like sclerite




Hemyda R-D. (1/1)

aurata R-D.



204.   Hind margin of eye straight or evenly convex. Ocellar triangle normal in shape, not dividing frontal vitta







205.   Palpus present. Vibrissae arising distinctly above lower margin of face. Legs yellowish basally, darkened apically


Xanthomelanodes 2nd Townsend (3/5)



205.   Palpus absent. Vibrissae level with lower margin of face. Legs entirely dark


Cylindromyia Meigen (11/17 in 5 subgenera)





206.   M ending in R4+5 well before wing margin






206.   M ending in wing margin or in R4+5 near margin








207.   Ocellar bristles lateroclinate or slightly reclinate. Calypter usually white, exceptionally large, extending beyond middle of syntergite1+2. Apex of female abdomen with pincers-like cerci, each with sharp apex and peg-like setae on inner margin





Leucostoma Meigen (5/9)



207.   Ocellar bristles proclinate or absent. Calypter not extending back to middle of syntergite1+2. Apex of female without pincers-like cerci (except in Clairvillia)







208.   Wing slender, usually darkened with whitish apex. M beyond bend meeting R4+5 at right angles; R4+5 + M twice as long as portion of M beyond bend



Catharosia Rondani (3/5)



208.   Wing not as described above; M not so abruptly bent anteriorly







209.   Wing patterned with brown spots, bars, and triangles






209.   Wing transparent, unpatterned







210.   Face with prominent oval central tubercle or carina. Antenna long, cylindrical; pedicel and first flagellomere each as long as arista. Lunule without setae. Thorax black. Apex of cell R4+5 dark




Euthera 2nd Loew (2/3)



 210.   Face with at most a low indistinct keel-like carina between antennae. Antennae short; length of pedicel and first flagellomere together no longer than arista. Lunule, in some species. With setulae. Thorax yellow or pale brown. Apex of cell R4+5 transparent



O. clausa


Oestrophasia B. & B. (3/4 in 2 subgenera)







211.   Proboscis extremely long and slender, curving forward, longer than height of head. Face pruinose, with thickened flat-topped carina separating very short antennae. Tergite 7 of female divided medially into two spine-covered lobes



Imitomyia Townsend (1/1)

sugens Loew



211.   Proboscis not exceptionally long, shorter than height of head. Face without a prominent carina. Tergite 7 of female without spine-covered lobes







212.   Eye very large, occupying almost all of side of head; gena reduced to a narrow band less than one-tenth height of head


S. triangulifera


Strongygaster Macquart, in part (3/4)



212.   Gena much wider, one-sixth height of head and more







213.   Eye covered with conspicuous dense hair. Facial ridge with setae on lower two-thirds or more; the uppermost of these setae situated at level of lowest frontal bristles; frontal bristles extending ventrally below level of base of arista


M. fracida


Mactomyia Reinhard (1/1)

fracida Reinhard



213.   Eye bare. Setae of facial ridge, if present, below level of lowest frontal bristle; frontal bristles not descending below level of base of arista


V. atrophopodoides


Vanderwulpia Townsend (1/2)

sequens Townsend




214.   Eye almost entirely covered with long hairs






214.   Eye apparently bare







215.   Ocellar bristles posterolaterally inclined; facial ridge with row of stout erect evenly spaced bristles. Proepisternum with pale hairs



C. semiotisthae                                                                            


Campylochaeta R-D. (9/11)



215.   Ocellar bristles proclinate or absent; facial ridge bare or with recumbent setae on lower half. Proepisternum bare







216.   Palpus reduced, parallel-sided, usually shorter than pedicel


L. tessellata


Linnaemya R-D., in part (4/7 in 2 subgenera)



216.   Palpus clavate, normal in length, longer than pedicel







217.   Body and most of head metallic green or blue


Gymnocheta R-D. (2/4)



217.   Body and head yellowish brown or black, not metallic green







218.   Middle basal bristle of postpronotum displaced anterolaterally, arising between anterior bristle and outer basal bristle or lateral to a line between them, and forming a nearly right-angled triangle with outer and inner basal bristles






218.   Middle basal bristle of postpronotum in line with outer and inner basal bristles








219.   Face convex, visible in profile; lower margin protruding anterior to vibrissal angle


P. ampelus


Panzeria R-D. (16/29)



219.   Face flat or deeply concave; neither central part nor lower margin visible in profile






220.   Abdominal tergite 5 orange, contrasting with dark preceding tergites; tergites 3 and 4 each with two pair of discal bristles, with one pair anterior to the other



Hineomyia Townsend (1/1)

setigera Coquillett



220.   Abdominal tergites concolorous; tergites 3 and 4 each with one pair of discal bristles or none







221.   Large species, over 8mm, shiny black or dark reddish brown, with yellow upper and lower calypteres and wing base. Antenna black


Melanophrys Williston (1/2)

insolita Walker



221.   Small species, less than 6mm, grayish brown to blackish, pruinose, with white calypteres and wing base. Antenna orange


Hyalurgus B. & B. (2/2)




222.   Anepimeral bristle well-developed, extending back well beyond middle of lower calypter






222.   Anepimeral bristle shorter, not extending to middle of lower calypter







223.   Arista thickened nearly to apex






223.   Arista thickened on less than basal half, tapering before middle to long thread-like apex


Return to 76





224.   Frontal bristles extending ventrally, in an evenly spaced row, nearly to lower margin of parafacial. First flagellomere of male sometimes divided longitudinally into two or three branches


D. robusta


Dichocera Williston (2/6)



224.   Frontal bristles extending ventrally only to level of base of first flagellomere; parafacial with a few hairs below lowest frontal bristle. Antenna of male undivided


O. aristalis


Ostracophyto Townsend (1/2)

flavicaudalis O'Hara





225.   Middorsal depression on abdominal syntergite 1+2 extending back to hind margin of syntergite






225.   Middorsal depression scarcely extending back beyond middle of syntergite







226.   Anterior and posterior lappets of posterior thoracic spiracle nearly equal in size; each lappet composed of a row of plumose hair-like extensions. Lower margin of face not protruding beyond vibrissal angle



Uramya R-D. (3/7)



226.   Anterior and posterior lappets unequal in size; posterior lappet subcircular. Lower margin of face protruding well beyond vibrissal angle. Arista slender. Palpus clubbed apically




Eriothrix Meigen (1/1)

penitalis Coquillett





227.   Base of R4+5 without setae, dorsally or ventrally. Scutellum with lateral, subapical, and apical bristles all more or less equivalent in size; apical bristles parallel or divergent. Antennal axis of head 1.5 times that of vibrissal axis



                                                                      C. bigelowi


Cleonice R-D. (2/3)



227.   Base of R4+5 with setae both dorsally or ventrally. Scutellum with stout divergent lateral bristles, short weak subapical bristles, and stout crossed apical bristles. Antennal axis of head less than 1.5 times that of vibrissal axis







228.   Genal groove relatively large, triangular, formed by sparation of parafacial and genal dilation from vibrissal angle. Eye height about two-thirds height of head. Three postsutural intra-alar bristles present


M. obscura


Macquartia R-D., in part (2/7)



228.   Genal groove nearly obliterated by the meeting of genal dilation, parafacial, and vibrissal angle. Eye height about four-fifths height of head. Two postsutural intra-alar bristles present


D. americana


Dufouria R-D. (1/1)

americana Reinhard




229.   Palpus minute, shorter than pedicel or absent





229.   Palpus well-developed, longer than pedicel







230.   Hind margin of eye indented. Prementum slender, elongate, more than twice as long as height of head. Abdomen orange red, slender, contrasting with dark wings


B. aelops


Beskia B. & B. (1/1)

aelops Walker



230.   Hind margin of eye straight or evenly convex. Prementum shorter than height of head. Abdomen and wings not as above







231.   Large species, 10 mm or more in length. Head and body mostly yellowish or pale brownish pruinescent. Abdomen slender, pointed apically. Apical scutellar bristles about half as long as subapical scutellar bristles; lateral scutellar bristles undeveloped


Icelia R-D. (1/1)

triquetra Olivier



231.   Smaller species, under 7 mm in length. Head and body mostly black or gray. Abdomen black. Apical scutellar bristles as long as lateral scutellar bristles; subapical scutellar bristles reduced








232.   Abdominal sternites unsclerotized; edges of tergites widely separated midventrally, connected by pale membrane, with terminalia tucked into resulting cavity; intersegmental junction between tergites 4 and 5 more prominent than between tergites 2 and 3 or tergites 3 and 4. Cell R4+5 opening at apex of wing




Besseria R-D., in part (2/3)



232.   Abdominal sternites fully sclerotized; connecting edges of tergites with little or no membrane exposed; intersegmental junctions between tergites 2-5 all equally will-defined. Cell R4+5 ending before apex of wing


Eloceria R-D. (1/2)

nigra Coquillett





233.   Ocellar setae inclined laterally or posterolaterally



Clairvillia R-D. (1/3)

timberlakei Walton



233.   Ocellar setae proclinate or absent







234.   Medial surface of fore coxa entirely covered with appressed setulae



Thelaira R-D. (1/2)

americana Brooks



234.   Medial surface of fore coxa bare; appressed setulae confined to anteromedial surface with bare area medially








235.   Lower proepimeral bristle well-developed, directed anteroventrally, in opposite direction to upper proepimeral bristle



Neaera R-D. (3/3)



235.   Lower proepimeral bristle absent, or reduced in size and directed dorsally, parallel to upper proepimeral








236.   M not reaching wing margin, ending about where bend should be


Besseria R-D., in part (2/3)



236.   M reaching wing margin, with bend in usual position







237.   Vibrissa not distinctly differentiated; genal groove exceptionally large; postgenal margin rather straight and vertical, ending below eye


E. exilis


Eutrixa Coquillett (1/2)

exilis Coquillett



237.   Vibrissa distinctly differentiated; postgenal margin extending forward below eye forming a distinct genal dilation









238.   Prosternum inflated



Ormia R-D. (8/9)



238.   Prosternum normal







239.   Face convex, visible in lateral view, at least on lower half






239.   Face concave at least on upper half, not visible in profile except sometimes at vibrissal angle







240.   Mid tibia with only one anterodorsal bristle or none






240.   Mid tibia with two or more anterodorsal bristles








241.   Pleuron with dense tufts of long white plumose hairs



Leskiopsis Townsend (1/1)

thecata Coquillett



241.   Pleuron hairs not plumose and not especially long or dense







242.   R4+5 setose more than halfway to crossveing r-m. Postpronotum with a small anterolateral bristle in addition to the basal bristles




Genea Rondani (7/7 in 2 subgenera)



242.   R4+5 with a few setae at base only. Postpronotum lacking an anterolateral bristle








243.   Mid tibia with one anterodorsal bristle


Leskia R-D. (2/3)



243.   Mid tibia without an anterodorsal bristle


Drepanoglossa Townsend (1/2)

tenuirostris Reinhard





244.   Arista thickened on basal two-thirds or more, tapering abruptly to slender apex. Prementum longer than fore femur



Aphria R-D. (2/2)



244.   Arista thickened on less than basal half, tapering gradually and uniformly to apex. Prementum shorter than fore femur



Solieria R-D. (1/5)

flava Townsend




245.   Prementum long and slender, often curved posteriorly, at least as long as height of eye, and in most species longer than height of head; labella minute, pointed, scarcely wider than apex of prementum






245.   Prementum shorter than height of eye; labella normal, pad-like







246.   Posterior margin of eye slightly concave (see image above). Antenna arising near level of middle of eye; first flagellomere no longer than half height of eye. Scutellum with well-developed lateral bristles



Epigrimyia Townsend (2/2)



246.   Posterior margin of eye convex. Antenna arising near upper margin of eye; first flagellomere large in both sexes, as long as height of eye. Scutellum lacking lateral bristles









247.   Membrane between lower facial margin and clypeus with a pair of convex subtriangular sclerites. Aristomere 2 elongate, two or more times as long as wide; first flagellomere truncate, widest beyond middle



                                                                       C. politura


Clausicella Rondani (6/8)



247.   Membrane between lower facial margin and clypeus without above mentioned sclerites. Aristomere 2 minute, no longer than wide; first flagellomere rounded apically, widest before middle



Crocinosoma Reinhard (1/1)

cornuale Reinhard





248.   Arista plumose, with the longest hairs longer than greatest width of arista







248.   Arista bare, or nearly so, with the longest hairs shorter than greatest width of arista







249.   Aristal hairs much longer at base of arista than at apex, arising all around arista. Parafacial with minute pale hairs. Midleg of male about twice as long as foreleg or hindleg



Cholomyia Bigot (1/1)

inaequipes Bigot



249.   Aristal hairs as long at apex of arista as on base (see images at 248). Parafacial bare. Midleg of male not exceptionally long







250.   Genal dilation extending anteriorly to vibrissal angle and parafacial, reducing genal groove to a sulcus. Small shiny brown species with little pale pruinosity



Anthomyiopsis Townsend (1/1)

cypseloides Townsend



250.   Genal dilation separated from vibrissal angle and parafacial by a distinct triangular pruinose genal groove. Species with extensive pale pruinose areas







251.   Scutellum with well-developed lateral bristles in addition to basal and apical bristles (see below, except that lateral bristles are parallel or convergent). R4+5 setose at least to crossvein r-m. Facial ridge usually with a few pale hairs above vibrissa. Female with sword-like ovipositor protruding from apex of abdomen, except in elegans (Reinhard)




Spathidexia Townsend (6/7 in 2 subgenera)



251.   Scutellum with well-developed basal and apical bristles only; lateral setae hair-like. R4+5 with one bristle at base. Facial ridge without pale hairs. Ovipositor tubular



Chaetonopsis Townsend (1/1)

spinosa Coquillett




252.   Scutum with only one pair of presutural acrostichal bristles, the posteriormost pair, which is situated anterior to scutellum





252.   Scutum usually with two pairs of presutural and two pairs of postsutural acrostichal bristles







253.   Scutellum without well-developed lateral bristles. Back of head convex


Euclytia Townsend (1/1)

flava Townsend



253.   Scutellum with well-developed lateral bristles as well as basal and apical bristles. Upper third or more of back of head concave



Opesia R-D. (1/1)

americana Bigot





254.   Eye, viewed in profile, occupying almost entire side of head, more than half as long (front to back) as high, thus reducing parafacial and gena to narrow bands



Strongygaster Macquart, in part (3/4)



254.   Eye, viewed in profile, less than half as long (front to back) as high; parafacial, gena and postgena more extensive








255.   Lateral scutellar bristles lacking



Phyllophilopsis Townsend, in part (2/2)



255.   Lateral scutellar bristles well-developed








256.   First flagellomere with sharply pointed apex. Scutellum short and broadly rounded, with crossed apical bristles and divergent lateral bristles arising rather far back between lateral and subapical position




Acemya R-D. (5/5)



256.   First flagellomere rounded apically. Scutellum more triangular with truncate apex; subapical scutellar bristles long and divergent, arising from corners of truncate apex








257.   Mid tibia without a ventral bristle. Abdomen of female arched ventrally, with telescopic ovipositor directed ventrally



Trigonospila Pokorny, in part (3/3)



257.   Mid tibia with well-developed ventral bristle. Abdomen of female not modified and ovipositor directed posteriorly







258.   Frons with row of proclinate orbital setae extending anteroventrally to level of lowest frontal bristle



Euhalidaya Walton, in part (1/1)

genalis Coquillett


258.   Frons with not more than two proclinate orbital bristles







259.   Facial ridge thickened, protruding beyond parafacial, thus visible in profile, usually with small appressed setulae on lower half or more. Arista, when orange, thickened nearly to apex



Phasmophaga Townsend, in part (3/4)



259.   Facial ridge narrow, without setae. Arista thickened only at base, tapering to long slender apex







260.   Katepisternum with two bristles


Anisia Wulp, in part (4/4)



260.   Katepisternum with three bristles


Opsomeigenia Townsend, in part  (2/) (includes Euthelyconychia) (3/4)