Key to the genera of Tachinidae

in the eastern US


Third page of the Tachinid key. See page one for information and initial key points. Eastern species list.

Morphology site of O'Hara


131.   R4+5 setose from base to well beyond crossvein r-m

 

 

132

 

 

131.   R4+5 not setose beyond crossvein r-m

 

 

133

 

 


 

 

132.   Katepisternum with a row of fine hairs ascending from midventral angle of katepisternum to a point just ventral to lower katepisternal bristle. Upper part of anepisternum usually with two well-developed bristles. Parafacial usually without minute setae below lowest frontal bristle

 

Actia R-D. (5/8)

 

 

132.   Katepisternum lacking such a row of fine hairs. Upper part of anepisternum usually with a single well-developed bristles. Parafacial with minute setae extending from lowest frontal bristle usually to level of base of arista

 

Ceromya R-D. (8/9)

 

 


 

 

133.   Scutellum lacking both lateral and discal bristles. Membrane between lower genal margin and clypeus forming a convex sclerite

 

 

 

Ginglymia Townsend (2/2)

 

 

133.   Scutellum with small lateral bristles and well-developed discal bristles. No sclerite between genal margin and clypeus

 

 

134

 

 


 

 

134.   R1 setose dorsally

 

 

Miamimyia Townsend (1/1)

cincta Townsend

 

 

134.   R1 bare

 

Chaetonodexodes Townsend, in part (1/1)

vanderwulpi Townsend

 

 


 

135.   Hind coxa with one or more hairs on posterior margin

 

136

 

 

135.   Hind coxa bare on posterior margin

 

 

144

 

 


 

 

136.   Parafacial with one or two bristles; these distinctly larger than surrounding hairs and situated near lower margin of eye

 

 

137

 

 

136.   Parafacial with hairs only or bare

 

139

 

 


 

 

137.   Palpus present, filiform, not enlarged apically

 

P. anaxias

 

Peleteria 2nd R-D. (4/39 in 4 subgenera)

 

 

137.   Palpus absent

 

138

 

 


 

 

138.   Parafacial with two stout bristles; these more or less equal in size, separated from each other by about half distance between upper parafacial bristles and lowest frontal bristle. Frontal bristles arising in two rows, the lowest bristle at level of lower fourth of eye

 

 

Deopalpus Townsend (2/11)

 

 

138.   Parafacial usually with a single stout bristle, sometimes with a smaller bristle immediately below it. Frontal bristles arising in a single row, the lowest bristle at level of middle of eye

 

 

Copecrypta Townsend (1/1)

ruficauda Wulp

 

 


 

139.   Palpus well-developed

 

140

 

 

139.   Palpus absent or reduced to a minute tubercle; abdomen black except for a triangular spot of white or gold pollen on tergite 5; tergite 3 with two or three pairs of median marginal bristles; tergite 4 with group of discal bristles

 

 

Epalpus Rondani (1/3)

signifer Walker

 

 


 

 

140.   Eye covered with conspicuous dense hairs

 

Jurinia R-D. (2/2)

 

 

140.   Eye apparently bare

 

141

 

 


 

 

141.   Abdominal tergite 3 with only one pair of median marginal bristles

 

142

 

 

141.   Abdominal tergite 3 with more than one pair (usually three or more pairs) of median marginal bristles

 

143

 

 


 

 

142.   First flagellomere, even in male, small, subcircular, bluntly rounded or truncate apically, always shorter than pedicel

 

 

Tachina Meigen (12/38 in 2 subgenera)

 

 

142.   First flagellomere kidney-shaped, somewhat pointed apically, in most species as long as, or longer than, pedicel, even in female; if first flagellomere shorter than pedicel (some females), then abdominal tergites 2-4 bluish pruinose, contrasting with nonpruinose tergite 5

 

 

Archytas Jaennicke (12/15 in 2 subgenera)

 

 


 

 

143.   Abdominal tergites 3 and 4 without median discal bristles; tergite 3 with three to five pairs of short stout median marginal bristles; the longest of these no more than twice as long as the shortest (see image at 141)

 

Juriniopsis Townsend (2/4)

 

 

143.   Abdominal tergites 3 and 4 each with several pairs of discal bristles forming a median patch or irregular transverse row; tergite 3 with a single pair of median marginal bristles and several much shorter ones less than a third their length. Base of wing and a small area around r-m darkened. Abdomen dark brown

 

 

Pararchytas B. & B. (2/3)

 

 


 

144.   All four of the following characters present simultaneously: only one frontal bristle arising below upper margin of pedicel; pedicel with one or two exceptionally long slender setae which are more than half as long as arista; arista pubescent or plumose; subvibrissal ridge with five or more slender bristles usually continuous with bristles of subcranial margin. Male with upper frontal bristles proclinate

 

 

145

 

 

144.   Differing from the above in the following: frontal bristles usually descending below level of pedicel, but if not, then either setae on pedicel all shorter than half aristal length or arista minutely pubescent or apparently bare; subvibrissal ridge usually with fewer than five bristles, usually not continuous with bristles along subvibrissal ridge. Male with upper frontal bristles reclinate

 

 

163

 

 


 

 

145.   Proepisternum setose

 

 

146

 

 

145.   Proepisternum bare, rarely with a single hair

 

 

154

 

 


 

 

146.   Proboscis long, slender, anteriorly curved at apex, more than twice as long as head height

 

 

Prosenoides B. & B. (2/3)

 

 

146.   Proboscis at most 1.5 times as long as head height

147

 

 


 

 

147.   Middorsal depression on abdominal syntergite 1+2 confined to anterior half or less of that segment. Median marginal bristles on this syntergite normally absent

 

 

Dinera R-D. (1/1)

grisescens Fallen

 

 

147.   Middorsal depression on abdominal syntergite 1+2 extending back to hind margin of that segment, or if not, then one pair of erect median marginal bristles present

148

 

 


 

 

148.   Integument and pruinescence entirely pale orange brown. Proboscis and palpus both exceptionally short, shorter than arista. Lower half of face, in anterior view, high and narrow; vibrissa arising above lower facial margin

 

P. flavus

 

Phasiops Coquillett (1/1)

flavus Coquillett

 

 

148.   Integument dark brown or black, with grayish pruinescence. Proboscis well-developed. Palpus either normally developed or absent. Lower half of face not narrowed, and vibrissa arising at level of lower facial margin

 

 

149

 

 


 

 

149.   Middorsal depression on abdominal syntergite 1+2 confined to anterior two-thirds of that segment

 

 

Phalacrophyto Townsend (1/1)

sarcophagina Coquillett

 

 

149.   Middorsal depression on abdominal syntergite 1+2 reaching to or nearly to hind margin of that segment

 

 

150

 

 


 

 

150.   Face with at most a small carina that scarcely separates antennae (see second image at 148)

 

Billaea R-D. (7/9)

 

 

150.   Face with a prominent median carina, usually high enough to separate the antennae along their entire length

 

 

151

 

 


 

 

151.   Proboscis slender, straight, about as long as height of head. Hind tibia of male with comb-like dorsolateral row of short evenly and closely set setae

 

 

Opsotheresia Townsend (2/2)

 

 

151.   Proboscis stouter, shorter than height of head. Hind tibia of male without comb-like dorsolateral row of setae

 

152

 

 


 

 

152.   Facial carina widest and most prominent ventrally, below level of apices of antennae

 

Dexia Meigen (1/1)

ventralis Aldrich

 

 

152.   Facial carina widest and most prominent near middle of face, between antennae

 

153

 

 


 

 

153.   Facial carina of rather uniform height, flattened along its crest, in anterior view widest near its midpoint, just below level of base of arista. Inner vertical bristles parallel or divergent. Anterior genal margin straight or slightly concave, ending ventrally below eye well behind its midpoint

 

 

Nicephorus Reinhard (1/1)

floridensis Reinhard

 

 

153.   Facial carina more conical, most prominent at level of junction of pedicel and first flagellomere, with rounded crest (see image at 150). Apices of inner vertical bristles convergent. Anterior genal margin convex, sometimes with a few small hairs extending across membrane of genal groove to lower corner of face

 

Ateloglossa Coquillett (8/14)

 

 


 

154.   Middorsal depression on abdominal syntergite 1+2 extending back to hind margin of that segment

 

 

155

 

 

154.   Middorsal depression of abdominal syntergite 1+2 smaller and shallower, not extending back to hind margin

 

 

158

 

 


 

 

155.   Proboscis long and slender, longer than height of head

 

156

 

 

155.   Proboscis shorter and stouter, not as long as head height

 

157

 

 


 

 

156.   Face with a broad, flat-topped carina, completely separating antennae (see image at 150)

 

Prosena Lepeletier & Serville (1/1)

siberita Fabricius

 

 

156.   Face without a median carina

 

Mochlosoma B. & B. (2/5)

 

 


 

 

157.   Fronto-orbital plate outside of row of frontal bristles almost devoid of setae except near ocellar triangle. Abdomen strongly pointed apically, especially in male

 

 

Zelia R-D. (6/12)

 

 

157.   Fronto-orbital plate with setae outside of frontal bristles extending forward to below middle of eye. Abdomen rounded apically, usually with a variegated pruinose pattern. Moderate sized species, about 10-15 mm.

 

Ptilodexia B. & B. (10/22)

 

 

 


 

 

158.   M ending in R4+5 before reaching wing margin

 

159

 

 

158.   M ending separately in wing margin

 

160

 

 


 

 

159.   Fused portion of R4+5 and M shorter than crossvein r-m (first wing below); second costal portion (between apices of Sc and R1) three or more times as long as costal bristle (second wing below)

 

 

 

Estheria R-D., in part (1/1)

cinerea Townsend

 

 

159.   Fused portion of R4+5 and M longer than crossvein r-m; second costal portion less than twice as long as costal bristle

 

 

Microchaetina Wulp, in part (2/8)

 

 


 

 

160.   Proboscis and palpus both exceptionally short, shorter than arista. Lower part of face, at and below level of vibrissae, narrowed, not protruding. First flagellomere no longer than pedicel

 

 

Megapariopsis Townsend (1/1)

opaca Coquillett

 

 

160.   Proboscis and palpus not so modified. Lower part of face not strongly narrowed. First flagellomere distinctly longer than pedicel

 

161

 

 


 

 

161.   Abdominal syntergite 1+2 elongate and strongly constricted to form a cylindrical petiole

 

Cordyligaster Macquart (1/1)

septentrionalis Townsend

 

 

161.   Base of abdomen not petiolate

 

162

 

 


 

 

162.   Wing strongly patterned, with the veins emphasized with dark brown. Body predominantly yellow. Abdominal syntergite 1+2 without marginal bristles

 

E. litturata

 

Euantha Wulp (1/1)

litturata Olivier

 

 

162.   Wing unpatterned. Body blue gray. Abdominal syntergite 1+2 with one pair of median marginal bristles

 

Ochrocera Townsend (1/1)

vaginalis Townsend

 

 


 

163.   Scutellum with one to three straight nearly erect preapical bristles arising on dorsal surface just anterior to apex. Anterior margin of postgena with at most a slight genal dilation. Last section of CuA1 (beyond crossvein dm-cu) between half as long and twice as long as preceding section

 

 

 

 

 

 

164

 

 

163.   Scutellum without a group of one to three straight preapical bristles. Anterior margin of postgena usually with well-developed genal dilation. Last section of CuA1 usually less than half as long as preceding section

 

 

174

 

 


 

 

164.   Anepimeron with an exceptionally long stout bristle, extending beyond apex of lower calypter

 

 

165

 

 

164.    Anepimeron with at most a short bristle not extending beyond middle of lower calypter

 

 

166

 

 


 

 

165.   Parafacial with two or three long proclinate bristles. R1 and CuA1 without setae dorsally

 

A. cinerea

 

Athrycia R-D. (1/1)

cinerea Coquillett

 

 

165.   Parafacial with only a few short setae below lowest frontal bristle. R1, R4+5, and CuA1 setose dorsally

 

H. cauta

 

 

Hypovoria Villeneuve (2/2)

 

 


 

 

166.   M ending in R4+5 before reaching wing margin

 

 

167

 

 

166.   M ending separately in wing margin

 

 

168

 

 


 

 

167.   Abdominal tergite 4, and usually 3 also, with median marginal bristles

 

Wagneria R-D. (4/5)

 

 

167.   Abdominal tergites 3 and 4 each lacking differentiated marginal and discal bristles

 

Kirbya R-D. (2/5 in 3 subgenera)

 

 


 

 

168.   Parafacial with a single large proclinate bristle arising just below lowest frontal bristle

 

V. ruralis

 

Voria R-D. (2/2)

 

 

168.   Parafacial with a row of proclinate bristles or patch of small decumbent setae

 

C. coquilletti

 

169

 

 


 

 

169.   Eye covered with long dense hairs (see above image)

 

Cyrtophleba Rondani (2/3)

 

 

169.   Eye apparently bare

 

170

 

 


 

 

170.   Sc with bristles ventrally; R1 usually setose ventrally as well as dorsally, at least at apex

 

C. atripennis

 

Chaetoplagia Coquillett (1/1)

atripennis Coquillett

 

 

170.   Sc and R1 bare ventrally

 

171

 

 


 

 

171.   Parafacial setae small and short, more or less uniform in length, none distinctly bristle-like

 

 

172

 

 

171.   Parafacial setae composed of bristles and hairs, differing distinctly in size, the bristles usually arranged in a vertical row

 

E. erucicola

 

173

 

 


 

 

172.   Prementum slender, straight and as long as or longer than height of head; labella reduced, pointed apically

 

P. similis

 

Plagiomima B. & B. (5/10)

 

 

172.   Prementum shorter than height of head; labella usually enlarged

 

Metaplagia Coquillett (3/6)

 

 


 

 

173.   Second aristomere elongate, more than three times as long as wide. Face (see second image at 171)

 

 

Euptilopareia Townsend (1/2)

erucicola Coquillett

 

 

173.   Second aristomere not more than twice as long as wide

 

Meleterus Aldrich (1/2)

nuperus Reinhard

 

 


 

174.   Parafacial setose on lower half or more

 

 

175

 

 

174.   Parafacial bare at least on lower half

 

198

 

 


 

 

175.   M ending in R4+5 before reaching wing margin; the united petiole at least half as long as that part of M beyond bend

 

 

176

 

 

175.   M ending separately in wing margin or in R4+5 very close to margin

 

 

185

 

 


 

 

176.   Abdominal tergites 3 and 4 lacking differentiated marginal and discal bristles. Small shining black species with fronto-orbital plate and all of parafacial shining black contrasting with thickened orange brown arista and antennal base

 

E. dacotensis

 

Euscopolia Townsend (1/1)

dakotensis Townsend

 

 

176.   Abdominal tergite 4, and usually 3 also, with median marginal bristles. Parafacial usually pruinescent

 

177

 

 


 

 

177.   Ocellar bristles reclinate. Antennal bases separated from one another. Calypter exceptionally large, white, extending over basal third of abdomen. Apex of female abdomen with pincers-like cerci opposing one another horizontally, each with sharp apex and inner margin armed with peg-like setae

 

 

 

 

                                                                     L. simplex

 

Leucostoma Meigen, in part (5/9)

 

 

177.   Ocellar bristles proclinate or absent. Antennal bases contiguous. Calypter not exceptionally large. Female abdomen without modified pincer-like cerci

 

178

 

 


 

 

178.   Second aristomere three or more times as long as wide

 

 

179

 

 

178.   Second aristomere scarcely longer than wide

 

181

 

 


 

 

179.   First aristomere nearly as long as the second, both of them four or five times as long as wide (Fig. 89). Lower facial margin protruding beyond bases of vibrissae. Acrostichal bristles absent or hair-like

 

P. stylata

 

Periscepsia Gistel, in part (3/8 in 2 subgenera)

 

 

179.   First aristomere less than half as long as the second. Lower facial margin not protruding. Acrostichal bristles well-developed

 

180

 

 


 

 

180.   Anepimeron bare, except for a single large bristle accompanied by one or two smaller ones at its base

 

Mauromyia Coquillett (4/5)

 

 

180.   Anepimeron with a group of small bristles

 

Periscepsia Gistel, in part (3/8 in 2 subgenera)

 

 


 

 

181.   Abdominal tergites 3 and 4 with median discal bristles

 

182

 

 

181.   Abdominal tergites 3 and 4 without discal bristles

 

184

 

 


 

 

182.   Prementum slender and elongate, 1.5 times head height, and curved forward; labella minute, appearing as a continuation of prementum. Abdominal tergite 3 with two pairs of discal bristles, on e ahead of the other and set far forward on the segment; hind margin of tergite 2 concave

 

T. skinneri

 

Trochiloides Coquillett (1/2)

leonardi West

 

 

182.   Prementum not longer than head height; labella enlarged. Abdominal tergite 3 with a single pair of discal bristles

183

 

 


 

 

183.   Parafacial setae few, small, often only one present, confined to lower fourth of parafacial

 

 

Loewia Egger (1/1)

foeda Meigen

 

 

183.   Parafacial setae long and erect, forming a row extending along entire length of parafacial

 

P. helymus

 

Periscepsia Gistel, in part (3/8 in 2 subgenera)

 

 


 

 

184.   Eye setose. Parafacial wider than width of first flagellomere in profile. Wing with long costal spine and dark anterior margin

 

Muscopteryx Townsend, in part (4/10)

 

 

184.   Eye bare. Parafacial narrower that width of first flagellomere. Wing with short costal bristle and scarcely any dark pigment

 

Paradidyma B. & B., in part (11/16)

 

 


 

185.   M gradually and evenly curved, without an abrupt bend. Parafacial forming a narrow band encircling lower margin of eye, ending at its posteroventral corner

 

F. montana

 

Freraea R-D. (1/1)

montana Coquillett

 

 

185.   M with a distinct bend. Parafacial not encircling lower margin of eye, ending before posteroventral part of eye

 

 

 

186

 

 


 

 

186.   Anterior border of postgena rather straight or concave, without pronounced genal dilation, meeting ventral cranial margin behind middle of head; genal groove thus rather extensive

 

 

187

 

 

186.   Anterior border of haired part of gena convex anteriorly, curving anteriorly to form a small to moderate genal dilation extending forward to a point anterior to middle of head; genal groove consequently much narrower

 

 

191

 

 


 

 

187.   Median surface of fore coxa almost entirely covered with short appressed black setae. Height of eye about half total height of head. Antenna small, recessed into small deep oval facial cavity. Lower part of face long and narrow, with four to five pairs of large subvibrissal bristles above lower facial margin

 

 

M. disjuncta

 

Microphthalma Macquart (3/4)

 

 

187.   Median surface of fore coxa with setae confined to anterolateral edge. Height of eye more than half total height of head. Antenna and face not highly modified as above

 

188

 

 


 

 

188.   Parafacial with single row of bristles

 

P. affinis

 

Paradidyma B. & B., in part (11/16)

 

 

188.   Parafacial irregularly covered with setae of varying sizes

 

189

 

 


 

 

189.   Parafacial setae usually of two sizes, namely a single row of larger bristles along anterior margin of parafacial sometimes even arising from membrane anterior to it, flanked by smaller and finer setae laterally. Integument of body of most species rather shiny, bluish of greenish black. Abdomen more than half as wide as long; tergite 6 and syntergosternite 7+8 exposed. Eye facets in male slightly larger on upper two-thirds of eye and when densely haired, hairs longer above than below

 

G. clistoides

 

Gnadochaeta Macquart (6/17)

 

 

189.   Parafacial setae not so differentiated into two series. More slender species with narrower gray pruinose, transversely banded abdomen; tergite 6 and syntergosternite 7+8 concealed beneath tergite 5. Eye facets not enlarged dorsally in male

 

190

 

 


 

 

190.   Anepimeron with one bristle extending back to hind margin of upper calypter. Female with tergite 6 longitudinally divided into two halves (see below). Male with unhinged aedeagus and well-developed gonopod

 

 

Pseudopachystylum Mik (1/1)

debile Townsend

 

 

190.   Anepimeron with hairs only. Female with tergite 6 undivided. Male with hinged aedeagus; gonopod reduced

 

 

Phyllomya R-D. (1/5)

polita Coquillett

 

 


 

191.   Middorsal depression of abdominal syntergite 1+2 extending back to hind margin of that segment; syntergite 1+2 lacking median marginal bristles

 

192

 

 

191.   Middorsal depression not extending back to hind margin of syntergite 1+2, usually not more than halfway back; syntergite 1+2 with median marginal bristles and sometimes with median discal bristles as well

 

195

 

 


 

 

192.   Palpus greatly reduced, parallel-sided, usually shorter than pedicel

 

L. tessallata

 

Linnaemya R-D. (4/7 in 2 subgenera)

 

 

192.   Palpus clavate, normal in length, longer than pedicel

 

193

 

 


 

 

193.   Bristles on lower part of parafacial reclinate, curved dorsally, forming an evenly spaced row that is continuous with frontal bristle row, and evidently an extension of it extending ventrally nearly to lower margin of parafacial. First flagellomere of male sometimes divided longitudinally into tow of three branches

 

D. robusta

 

Dichocera Williston (2/6)

 

 

193.   Bristles on lower part of parafacial proclinate, curved ventrally, separated from reclinate lowermost bristle or frontal row at level of insertion of arista

 

194

 

 


 

 

194.   Aristomeres 1 and 2 elongate, all three subequal in length. R1 and R4+5 each setose dorsally along most of their length

 

 

Triarthria Stevens (1/1)

setipennis Fallen

 

 

194.   Aristomeres 1 and 2 scarcely longer than wide, much shorter than aristomere 3. R1 bare and R4+5 setose only at base, or bare. Antennal bases contiguous or nearly so; antenna orange

 

Lypha R-D. (5/6)

 

 


 

195.   Parafacial with single evenly spaced row of long, stout setae, increasing in size ventrally, usually also with some shorter finer hairs laterally

 

 

196

 

 

195.   Parafacial with scattered setae not arranged in a row

 

197

 

 


 

 

196.   Aristomere 3 strongly swollen only on basal fourth or less, tapering abruptly to apex. First flagellomere no longer than 1.5 times length of pedicel. Vibrissa arising above level of lower facial margin

 

 

Muscopteryx Townsend, in part (4/10)

 

 

196.   Aristomere 3 thickened on basal third or more before tapering to apex. First flagellomere two or more times length of pedicel. Vibrissa arising at level of lower facial margin

 

Blepharomyia Townsend (1/2)

tibialis Curran